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Mayfly Genus Ephemerella (Hendricksons, Sulphurs, PMDs)

Taxonomic Navigation -?-
Kingdom Animalia (Animals)
» Phylum Arthropoda (Arthropods)
» Class Insecta (Insects)
» Order Ephemeroptera (Mayflies)
» Family Ephemerellidae (Hendricksons, Sulphurs, PMDs, BWOs)
» Genus Ephemerella (Hendricksons, Sulphurs, PMDs)
Species in EphemerellaNumber of SpecimensNumber of Pictures
Ephemerella aurivillii730
Ephemerella dorotheaSulphur211
Ephemerella excruciansPale Morning Dun15
Ephemerella invariaSulphur47202
Ephemerella needhamiLittle Dark Hendrickson1786
Ephemerella septentrionalis325
Ephemerella subvariaHendrickson34162

11 species aren't included.
Common Name
Pictures Below
Ephemerella used to be a "super-genus" containing a myriad of extremely important species, many of which are now spread out among other genera in the Ephemerellidae family. These include the Large Blue-Winged Olives of the Drunella genus and important species like Attenella attenuata, Dannella simplex, and Serratella deficiens. Despite these reclassifications, the Ephemerella genus still contains some of the most important species in North America.

There is a lot of variation between the species; read about each one for the details.
  

Where & When

Regions: East, Midwest, West
Time Of Year (?): Mostly in the spring in the East, summer in the West
Ephemerella nymphs inhabit all types of water. This genus is most important in the East and Midwest, where the major species hatch in the spring and a few continue into the summer. In the West, the other closely related genus Drunella takes on greater importance, but the Pale Morning Duns of Ephemerella are still important in the summer.

Spinner Behavior

The concentrated evening spinner falls of many species in this genus are some of the best fly fishing events of the spring.

They are also one of the groups of mayflies most prone to the unfortunate behavior of mating and ovipositing over blacktop roads hundreds of yards from the river.

122 Mayfly Specimens in the Genus Ephemerella:

Specimen Page:1234...13
Ephemerella invaria (Sulphur) Mayfly SpinnerEphemerella invaria (Sulphur) Mayfly SpinnerView 12 Pictures
Region: Upper Midwest
Collected Jun 3, 2005
Added May 25, 2006
Ephemerella needhami (Little Dark Hendrickson) Mayfly SpinnerEphemerella needhami (Little Dark Hendrickson) Mayfly SpinnerView 12 Pictures
Region: Upper Midwest
Collected Jun 4, 2005
Added May 25, 2006
Ephemerella septentrionalis Mayfly DunEphemerella septentrionalis  Mayfly DunView 10 PicturesI'm pretty sure this dun belongs to septentrionalis, because her legs are just too long for invaria. I know that species is in this system because I collected several nymphs some miles downstream.

This really pretty mayfly was in kind of bad shape when I found it crippled on the surface, and bouncing around in my container with a bunch of green drakes didn't help.
Region: Catskills
Collected Jun 1, 2007
Added Jun 4, 2007
Specimen Page:1234...13

6 Streamside Pictures of Ephemerella Mayflies:

Streamside Photo Page:12
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Here's an above-the-water view of a stillborn Ephemerella subvaria dun which I also photographed from below the water.
Here's an above-the-water view of a stillborn (
This stillborn Ephemerella subvaria dun is trapped in its shuck.
This stillborn Ephemerella subvaria dun is trapped in its shuck.
Stillborn: In fly fishing, a stillborn insect is one which got stuck in its nymphal or pupal shuck during emergence and floats helplessly on the surface instead of flying away. It is a specific class of cripple, although it is sometimes used interchangeably with that term.
)
Ephemerella subvaria dun which I also photographed from below the water.

In this picture: Mayfly Species Ephemerella subvaria (Hendrickson).
RegionUpper Midwest
Date TakenApr 19, 2006
Date AddedApr 23, 2006
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This Ephemerella invaria sulphur dun got stuck in its shuck trying to emerge.  This isn't exactly a "natural" pose for a photograph, but it kind of shows what an emerger pattern could look like.
This Ephemerella invaria sulphur dun got stuck in its shuck (
Here's an underwater view of the pupal shucks of several already-emerged Brachycentrus numerosus caddisflies.
Here's an underwater view of the pupal shucks of several already-emerged Brachycentrus numerosus caddisflies.
Shuck: The shed exoskeleton left over when an insect molts into its next stage or instar. Most often it describes the last nymphal or pupal skin exited during emergence into a winged adult.
)
trying to emerge. This isn't exactly a "natural" pose for a photograph, but it kind of shows what an emerger pattern could look like.

In this picture: Mayfly Species Ephemerella invaria (Sulphur).
RegionUpper Midwest
Date TakenMay 20, 2007
Date AddedJun 6, 2007
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RegionNortheast
Date TakenApr 30, 2007
Date AddedMay 3, 2007
Streamside Photo Page:12

37 Underwater Pictures of Ephemerella Mayflies:

Underwater Photo Page:12345
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This Ephemerella subvaria (Hendrickson) nymph picture is one of my favorites.
This Ephemerella subvaria (Hendrickson) nymph picture is one of my favorites.

In this picture: Mayfly Species Ephemerella subvaria (Hendrickson).
RegionUpper Midwest
Date TakenMar 24, 2004
Date AddedJan 25, 2006
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RegionUpper Midwest
Date TakenApr 23, 2004
Date AddedJan 25, 2006
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There's a large Ephemerella subvaria nymph in the top left.
RegionUpper Midwest
Date TakenMar 20, 2004
Date AddedJan 25, 2006
Underwater Photo Page:12345

Recent Discussions of Ephemerella

Aw Shucks 7 Replies »
Posted by Martinlf on May 19, 2009 in the species Ephemerella invaria
Last reply on May 21, 2009 by Wiflyfisher
OK, this is going to seem like a major duh experience for some of you, but the other night I found a sulphur spinner on the door of a bathhouse in a campground I was staying at. Looking for other bugs I then saw a pale nymph shuck on the door. I was totally confused. A nymph this far from the stream? Was this some alien bug? Looking closer I noticed that the shape was too slender for a nymph and that the wing pads were more like little protruding pockets--and it hit me. Spinner shuck. I knew that mayflies molted to produce a spinner, but I had thought the shuck would be more insubstantial--something that would be flimsy and lack form. This was so cool, and at the same time I felt so silly for thinking it could somehow have been a nymph shuck. It's the first spinner shuck I've seen, but I assume that I'll start seeing them everywhere now, like a new word you learn. Anybody else have a spinner shuck story?
Replypmd hatches in ny? 3 Replies »
Posted by Trouthunter on Apr 18, 2009
Last reply on Apr 19, 2009 by GONZO
I was recently told by a close friend of mine that in the southertier of ny ( finger lakes region) that we have very good hatches of pmds. I was hoping that someone could confirm this or otherwise before I tye up patterns. I have seen what I believe are sulphurs with slightly the same body and wing coloration but again thought that these couldn't be the famed insect.
Replypmd hatches in ny?
Posted by Trouthunter on Apr 18, 2009
I was recently told by a close friend of mine that in the southertier of ny ( finger lakes region) that we have very good hatches of pmds. I was hoping that someone could confirm this or otherwise before I tye up patterns. I have seen what I believe are sulphurs with slightly the same body and wing coloration but again thought that these couldn't be the famed insect.
ReplyHatching Hendrickson
Posted by Martinlf on Oct 14, 2008 in the species Ephemerella subvaria
Here's another hatching mayfly, this one stillborn. Click on "31 more specimens" and scroll down.
ReplyHatching Sulphur
Posted by Martinlf on Oct 14, 2008 in the species Ephemerella invaria
Click on "40 more specimens" and scroll down for one photo.
Reply
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