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Mayfly Species Ephemerella excrucians (Pale Morning Dun)

Taxonomic Navigation -?-
Kingdom Animalia (Animals)
» Phylum Arthropoda (Arthropods)
» Class Insecta (Insects)
» Order Ephemeroptera (Mayflies)
» Family Ephemerellidae (Hendricksons, Sulphurs, PMDs, BWOs)
» Genus Ephemerella (Hendricksons, Sulphurs, PMDs)
» Species excrucians (Pale Morning Dun)
Common Names
Pictures Below
Until recently, Ephemerella excrucians was a widespread species of relatively minor importance. Recent work by entomologists uncovered that it is the same species as the important Western Pale Morning Duns Ephemerella inermis, and the two combined took the name of excrucians.

The behavioral information I've given below is tentative. Because of the taxonomic shuffling and the fact that books usually describe this species together with others, it is a little bit unclear which writings specifically apply to excrucians.
  

Where & When

Regions: East, Midwest, West
Time Of Year (?): May to October in the West, peaking in July and August; May and June in the East
Preferred Waters: Mostly streams and large rivers, but sometimes Western mountain lakes
These mayflies are extremely abundant throughout the mountain West. Books give a wide range of dates for their emergence, so anglers should consult hatch charts for their specific rivers to time this species.

The former Western species Ephemerella lacustris is described in angler's literature as occasionally important in mountain lakes. Like inermis, it was discovered to be a synonym (Synonym: A former name of a taxon, usually a species. Entomologists frequently discover that two insects originally described as different species are one in the same, and they drop one of the names. The dropped name is said to be a synonym of the remaining name. These changes take a while to trickle into the common knowledge of anglers; for example, Baetis vagans is now a synonym of Baetis tricaudatus.) of excrucians.

Hatching Behavior

Time Of Day (?): Late morning and early evening in the West; late afternoon to evening in the East
Water Temperature: 55-60°F
Several authors have noted that trout are very selective to this species. The emergence puzzles mirror those of the closely related species Ephemerella dorothea.

The Western emergence may last for most of the day on prime rivers, and it commonly lasts 2-3 hours.

Spinner Behavior

Time Of Day: Morning and again at dusk in the West; only dusk in the East, where they're not important
Habitat: Riffles
The spinners are as important as the duns in the West. In the East, according to Knopp & Cormier, the spinner falls are "an insignificant event for the fly fisher."

Nymph Biology

Current Speed: Usually slow in the West; medium to fast in the East
Substrate: Many substrates, but gravel is best
In Mayflies: An Angler's Study of Trout Water Ephemeroptera , Knopp & Cormier write specifically of excrucians that it can be found in riffles and fast runs in the East. This contrasts with the slow-water preference of the ex-inermis mayflies in the West.

The nymphs can be important to imitate before and during the emergence. Swisher and Richards in Selective Trout note that the nymphs come in either olive or very dark brown colors and trout can be selective to one or the other.

1 Mayfly Specimen in the Species Ephemerella excrucians:

Ephemerella excrucians (Pale Morning Dun) Mayfly NymphEphemerella excrucians (Pale Morning Dun) Mayfly NymphView 5 PicturesI spent (Spent: The wing position of many aquatic insects when they fall on the water after mating. The wings of both sides lay flat on the water. The word may be used to describe insects with their wings in that position, as well as the position itself.) a while with a microscope to fairly positively identify this specimen as Ephemerella excrucians.
Region: Upper Midwest
Collected Jun 9, 2005
Added May 26, 2006

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