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Dark Red Quills

Like most common names, "Dark Red Quill" can refer to more than one taxon. They're previewed below, along with 11 specimens. For more detail click through to the scientific names.

Mayfly Genus Cinygmula

These are often called Dark Red Quills.
This is primarily a Western genus. Cinygmula ramaleyi is the most important species, producing good hatches in the West. Cinygmula reticulata may also be relevant.

There is only one Eastern species, Cinygmula subaequalis, and its importance is minor.
Cinygmula subaequalis (Small Gordon Quill) Mayfly NymphCinygmula subaequalis (Small Gordon Quill) Mayfly Nymph View 10 PicturesThis nymph is missing a few gills, but is otherwise in good shape. It was the only one of its species which turned up in my sample.
Collected May 29, 2007 from in
Added to Troutnut.com by on June 4, 2007
Female Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly DunFemale Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly Dun View 7 PicturesThis dun is almost certainly of the same species as this nymph, as it hatched in my cooler from a nearly identical nymph.
Collected July 10, 2011 from in
Added to Troutnut.com by on July 13, 2011
Male Cinygmula (Dark Red Quills) Mayfly SpinnerMale Cinygmula (Dark Red Quills) Mayfly Spinner View 11 PicturesI'm unsure of the ID on this one; keys put it closest to Cinygmula reticulata, but I'm very doubtful of the species and not positive on the genus. Epeorus is another possibility, but I don't know which species it would be.

This one was collected in association with a female dun probably of the same species.
Collected July 1, 2017 from in
Added to Troutnut.com by on July 2, 2017
Male Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly AdultMale Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly Adult View 4 PicturesAdults were collected from the North Fork of the Touchet River at Touchet Corral, 21 Sept. One photo is the swarm of males over the stream about 3 PM, air temp about 66 degree.
Collected September 21, 2011 from in
Added to Troutnut.com by on September 23, 2011

Mayfly Species Rhithrogena impersonata

These are sometimes called Dark Red Quills.
This intriguing species has two distinct colors of nymphs, which were once considered to be different species. Most nymphs are a dark olive gray, but some are a surprisingly bright reddish brown. The red ones were once classified as Rhithrogena sanguinea. There is no apparent difference between the adults of the two varieties.
Rhithrogena impersonata (Dark Red Quill) Mayfly NymphRhithrogena impersonata (Dark Red Quill) Mayfly Nymph View 6 PicturesThis was the only Rhithrogena specimen in a large sample of nymphs from a small Catskill stream. It looks virtually identical to Rhithrogena impersonata specimens collected in the Midwest, but I didn't get to check the distinguishing features under a microscope.
Collected April 19, 2006 from in
Added to Troutnut.com by on April 21, 2006

Mayfly Species Rhithrogena jejuna

These are sometimes called Dark Red Quills.

Mayfly Species Attenella attenuata

These are very rarely called Dark Red Quills.
This intriguing species has received a lot of attention in past angling books. Recent authors suspect that much of this credit was a case of mistaken identity, with Attenella attenuata receiving praise for the hatches of Drunella lata and Dannella simplex. Much of the credit was legitimate and accurate, but this species is no longer thought to be on par with its most popular cousins in Ephemerella and Drunella.

I have several specimens listed under this species, but I'm not positive the identification is correct.
Female Attenella attenuata (Small Eastern Blue-Winged Olive) Mayfly DunFemale Attenella attenuata (Small Eastern Blue-Winged Olive) Mayfly Dun View 4 PicturesThis specimen came from the same hatch as a male.
Collected June 8, 2005 from in
Added to Troutnut.com by on May 26, 2006

Mayfly Species Isonychia bicolor

These are very rarely called Dark Red Quills.
This is by far the most important species of Isonychia. Many angling books once split its credit with the species Isonychia sadleri and Isonychia harperi, but entomologists have since discovered that those are just variations of this abundant species.

See the main Isonychia page for more about these intriguing mayflies.
Isonychia bicolor (Mahogany Dun) Mayfly NymphIsonychia bicolor (Mahogany Dun) Mayfly Nymph View 7 PicturesThis Isonychia bicolor nymph from the Catskills displays the prominent white stripe sometimes characteristic of its species. This is the first such specimen I've photographed, because members of the same species in the Upper Midwest have a more subdued stripe (and were once thought to be a different species, Isonychia sadleri). The striking coloration on this eastern nymph is more appealing.
Collected April 19, 2006 from in
Added to Troutnut.com by on April 21, 2006
Female Isonychia bicolor (Mahogany Dun) Mayfly DunFemale Isonychia bicolor (Mahogany Dun) Mayfly Dun View 13 Pictures
Collected June 14, 2005 from in
Added to Troutnut.com by on May 26, 2006
Male Isonychia bicolor (Mahogany Dun) Mayfly SpinnerMale Isonychia bicolor (Mahogany Dun) Mayfly Spinner View 15 PicturesI got several really nice pictures of this spinner. I also collected a female on the same trip.
Collected August 9, 2006 from in
Added to Troutnut.com by on August 11, 2006

Mayfly Species Cinygmula ramaleyi

These are very rarely called Dark Red Quills.
This can be the first mayfly of the season on high mountain streams in the western states, but emerges later in the season in Alaska. It is the most important species of Cinygmula for anglers.
Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly NymphCinygmula ramaleyi (Small Western Gordon Quill) Mayfly Nymph View 9 PicturesThis nymph is almost definitely the same species as this dun, which hatched from a nearly identical nymph from the same collection.
Collected July 10, 2011 from in
Added to Troutnut.com by on July 13, 2011
Female Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly DunFemale Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly Dun View 7 PicturesThis dun is almost certainly of the same species as this nymph, as it hatched in my cooler from a nearly identical nymph.
Collected July 10, 2011 from in
Added to Troutnut.com by on July 13, 2011
Male Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly SpinnerMale Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly Spinner View 4 PicturesThis one was preserved in ethanol and partially dissected, so it doesn't have its natural wild coloration, and it's missing some appendages. I'm posting it for identification purposes.
Collected September 4, 2015 from in
Added to Troutnut.com by on October 27, 2015
Male Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly AdultMale Cinygmula ramaleyi (Small Western Gordon Quill) Mayfly Adult View 4 PicturesAdults were collected from the North Fork of the Touchet River at Touchet Corral, 21 Sept. One photo is the swarm of males over the stream about 3 PM, air temp about 66 degree.
Collected September 21, 2011 from in
Added to Troutnut.com by on September 23, 2011

Mayfly Species Rhithrogena undulata

These are very rarely called Dark Red Quills.
This is one of the two most common species of Rhithrogena.
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