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Dark Red Quills

Like most common names, "Dark Red Quill" can refer to more than one taxon. They're previewed below, along with 10 specimens. For more detail click through to the scientific names.

Mayfly Genus Cinygmula

These are often called Dark Red Quills.
This is primarily a Western genus. Cinygmula ramaleyi is the most important species, producing good hatches in the West. Cinygmula reticulata may also be relevant.

There is only one Eastern species, Cinygmula subaequalis, and its importance is minor.
Cinygmula subaequalis (Small Gordon Quill) Mayfly NymphCinygmula subaequalis (Small Gordon Quill) Mayfly NymphView 10 PicturesThis nymph is missing a few gills, but is otherwise in good shape. It was the only one of its species which turned up in my sample.
Region: Poconos
Collected May 29, 2007
Added Jun 4, 2007

Mayfly Species Rhithrogena impersonata

These are sometimes called Dark Red Quills.
This intriguing species has two distinct colors of nymphs, which were once considered to be different species. Most nymphs are a dark olive gray, but some are a surprisingly bright reddish brown. The red ones were once classified as Rhithrogena sanguinea. There is no apparent difference between the adults of the two varieties.
Rhithrogena impersonata (Dark Red Quill) Mayfly NymphRhithrogena impersonata (Dark Red Quill) Mayfly NymphView 8 PicturesThis specimen shows one of two distinct color types of Rhithrogena impersonata nymphs. This type is very reddish, especially in the gills, and it was formerly in its own species called Rhithrogena sanguinea.
Region: Upper Midwest
Collected Jun 5, 2005
Added May 25, 2006

Mayfly Species Rhithrogena jejuna

These are sometimes called Dark Red Quills.

Mayfly Species Attenella attenuata

These are very rarely called Dark Red Quills.
This intriguing species has received a lot of attention in past angling books. Recent authors suspect that much of this credit was a case of mistaken identity, with Attenella attenuata receiving praise for the hatches of Drunella lata and Dannella simplex. Much of the credit was legitimate and accurate, but this species is no longer thought to be on par with its most popular cousins in Ephemerella and Drunella.

I have several specimens listed under this species, but I'm not positive the identification is correct.
Female Attenella attenuata (Small Eastern Blue-Winged Olive) Mayfly DunFemale Attenella attenuata (Small Eastern Blue-Winged Olive) Mayfly DunView 4 PicturesThis specimen came from the same hatch as a male.
Region: Upper Midwest
Collected Jun 8, 2005
Added May 26, 2006
Female Attenella attenuata (Small Eastern Blue-Winged Olive) Mayfly SpinnerFemale Attenella attenuata (Small Eastern Blue-Winged Olive) Mayfly SpinnerView 11 Pictures
Region: Upper Midwest
Collected Jul 1, 2005
Added Apr 22, 2006

Mayfly Species Isonychia bicolor

These are very rarely called Dark Red Quills.
This is by far the most important species of Isonychia. Many angling books once split its credit with the species Isonychia sadleri and Isonychia harperi, but entomologists have since discovered that those are just variations of this abundant species.

See the main Isonychia page for more about these intriguing mayflies.
Isonychia bicolor (Mahogany Dun) Mayfly NymphIsonychia bicolor (Mahogany Dun) Mayfly NymphView 7 PicturesThis Isonychia bicolor nymph from the Catskills displays the prominent white stripe sometimes characteristic of its species. This is the first such specimen I've photographed, because members of the same species in the Upper Midwest have a more subdued stripe (and were once thought to be a different species, Isonychia sadleri). The striking coloration on this eastern nymph is more appealing.
Region: Catskills
Collected Apr 19, 2006
Added Apr 21, 2006
Female Isonychia bicolor (Mahogany Dun) Mayfly DunFemale Isonychia bicolor (Mahogany Dun) Mayfly DunView 13 Pictures
Region: Upper Midwest
Collected Jun 14, 2005
Added May 26, 2006
Male Isonychia bicolor (Mahogany Dun) Mayfly SpinnerMale Isonychia bicolor (Mahogany Dun) Mayfly SpinnerView 15 PicturesI got several really nice pictures of this spinner. I also collected a female on the same trip.
Region: Northeast
Collected Aug 9, 2006
Added Aug 11, 2006

Mayfly Species Ephemerella invaria

These are very rarely called Dark Red Quills.
This species, the primary "Sulphur" hatch, stirs many feelings in the angler. There is nostalgia for days when everything clicked and large, selective trout were brought to hand. There is the bewildering memory of towering clouds of spinners which promise great fishing and then vanish back into the aspens as night falls. There is frustration from the maddening selectivity with which trout approach the emerging duns--a vexing challenge that, for some of us, is the source of our excitement when Sulphur time rolls around.

Ephemerella invaria is one of the two species frequently known as Sulphurs (the other is Ephemerella dorothea). There used to be a third, Ephemerella rotunda, but entomologists recently discovered that invaria and rotunda are a single species with an incredible range of individual variation. This variation and the similarity to dorothea make matching this hatch exceptionally tricky.

As the combination of two already prolific species, this has become the most abundant of all mayfly species in Eastern and Midwestern trout streams.
Ephemerella invaria (Sulphur) Mayfly NymphEphemerella invaria (Sulphur) Mayfly NymphView 8 PicturesThis small Ephemerella invaria nymph was at least a month away from emergence.
Region: Catskills
Collected Apr 19, 2006
Added Apr 21, 2006
Female Ephemerella invaria (Sulphur) Mayfly DunFemale Ephemerella invaria (Sulphur) Mayfly DunView 6 Pictures
Region: Upper Midwest
Collected May 28, 2005
Added May 24, 2006
Male Ephemerella invaria (Sulphur) Mayfly SpinnerMale Ephemerella invaria (Sulphur) Mayfly SpinnerView 12 Pictures
Region: Upper Midwest
Collected Jun 3, 2005
Added May 25, 2006

Mayfly Species Cinygmula ramaleyi

These are very rarely called Dark Red Quills.
This can be the first mayfly of the season on high mountain streams. It is the most important species of Cinygmula.

Mayfly Species Rhithrogena undulata

These are very rarely called Dark Red Quills.
This is one of the two most common species of Rhithrogena.
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Last update July 19th, 2007.
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